US2003064499A1PendingUtilityA1

Hepatitis C virus protease

Priority: Apr 4, 1990Filed: Jun 18, 2001Published: Apr 3, 2003
Est. expiryApr 4, 2010(expired)· nominal 20-yr term from priority
C07K 2319/02C12N 2770/24222C07K 14/00C07K 2319/95C07K 2319/00C12N 9/0089A61K 38/00C12N 9/506C07K 14/005C12N 9/90
60
PatentIndex Score
0
Cited by
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References
0
Claims

Abstract

The protease necessary for polyprotein processing in Hepatitis C virus is identified, cloned, and expressed. Proteases, truncated protease, and altered proteases are disclosed which are useful for cleavage of specific polypeptides, and for assay and design of antiviral agents specific for HCV.

Claims

exact text as granted — not AI-modified
What is claimed:  
     
         1 . A composition comprising a purified proteolytic polypeptide derived from Hepatitis C virus.  
     
     
         2 . The composition of  claim 1 , wherein said polypeptide has a partial internal sequence substantially as follows: 
 Trp Thr Val Tyr His Gly Ala Gly Thr Arg Thr.    
     
     
         3 . The composition of  claim 1 , wherein said polypeptide has a partial internal sequence substantially as follows: 
 Leu Lys Gly Ser Ser Gly Gly Pro Leu.    
     
     
         4 . The composition of  claim 2 , wherein said polypeptide has substantially the partial internal sequence:  
       
         
           
                 
                 
               
                     
                 
                   Arg Arg Gly Arg Glu Ile Leu Leu Gly Pro Ala Asp Gly Met Val Ser Lys Gly 
                     
                 
                     
                 
                   Trp Arg Leu Leu Ala Pro Ile Thr Ala Tyr Ala Gln Gln Thr Arg Gly Leu Leu 
                 
                     
                 
                   Gly Cys Ile Ile Thr Ser Leu Thr Gly Arg Asp Lys Asn Gln Val Glu Gly Glu 
                 
                     
                 
                   Val Gln Ile Val Ser Thr Ala Ala Gln Thr Phe Leu Ala Thr Cys Ile Asn Gly 
                 
                     
                 
                   Val Cys Trp Thr Val Tyr His Gly Ala Gly Thr Arg Thr Ile Ala Ser Pro Lys 
                 
                     
                 
                   Gly Pro Val Ile Gln Met Tyr Thr Asn Val Asp Gln Asp Leu Val Gly Trp Pro 
                 
                     
                 
                   Ala Pro Gln Gly Ser Arg Ser Leu Thr Pro Cys Thr Cys Gly Ser Ser Asp Leu 
                 
                     
                 
                   Tyr Leu Val Thr Arg His Ala Asp Val Ile Pro Val Arg Arg Arg Gly Asp Ser 
                 
                     
                 
                   Arg Gly Ser Leu Leu Ser Pro Arg Pro Ile Ser Tyr Leu Lys Gly Ser Ser Gly 
                 
                     
                 
                   Gly Pro Leu Leu Cys Pro Ala Gly His Ala Val Gly Ile Phe Arg Ala Ala Val 
                 
                     
                 
                   Cys Thr Arg Gly Val Ala Lys Ala Val Asp Phe Ile Pro Val Glu Asn Leu Glu 
                 
                     
                 
                   Thr Thr Met Arg. 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         5 . The composition of  claim 1 , wherein said polypeptide has substantially the amino acid sequence shown in FIG. 1.  
     
     
         6 . A fusion protein, comprising: 
 a suitable fusion partner, fused to    a proteolytic polypeptide derived from Hepatitis C virus.    
     
     
         7 . The fusion protein of  claim 6 , wherein said fusion partner comprises human superoxide dismutase.  
     
     
         8 . The fusion protein of  claim 6 , wherein said proteolytic polypeptide has a partial internal sequence substantially as follows: 
 Trp Thr Val Tyr His Gly Ala Gly Thr Arg Thr.    
     
     
         9 . The fusion protein of  claim 6 , wherein said proteolytic polypeptide has a partial internal sequence substantially as follows: 
 Leu Lys Gly Ser Ser Gly Gly Pro Leu.    
     
     
         10 . The fusion protein of  claim 6 , wherein said proteolytic polypeptide has as a partial internal sequence:  
       
         
           
                 
                 
               
                     
                     
                 
                     
                   Arg Arg Gly Arg Glu Ile Leu Leu Gly Pro Ala Asp 
                 
                     
                     
                 
                     
                   Gly Met Val Ser Lys Gly Trp Arg Leu Leu Ala Pro 
                 
                     
                     
                 
                     
                   Ile Thr Ala Tyr Ala Gln Gln Thr Arg Gly Leu Leu 
                 
                     
                     
                 
                     
                   Gly Cys Ile Ile Thr Ser Leu Thr Gly Arg Asp Lys 
                 
                     
                     
                 
                     
                   Asn Gln Val Glu Gly Glu Val Gln Ile Val Ser Thr 
                 
                     
                     
                 
                     
                   Ala Ala Gln Thr Phe Leu Ala Thr Cys Ile Asn Gly 
                 
                     
                     
                 
                     
                   Val Cys Trp Thr Val Tyr His Gly Ala Gly Thr Arg 
                 
                     
                     
                 
                     
                   Thr Ile Ala Ser Pro Lys Gly Pro Val Ile Gln Met 
                 
                     
                     
                 
                     
                   Tyr Thr Asn Val Asp Gln Asp Leu Val Gly Trp Pro 
                 
                     
                     
                 
                     
                   Ala Pro Gln Gly Ser Arg Ser Leu Thr Pro Cys Thr 
                 
                     
                     
                 
                     
                   Cys Gly Ser Ser Asp Leu Tyr Leu Val Thr Arg His 
                 
                     
                     
                 
                     
                   Ala Asp Val Ile Pro Val Arg Arg Arg Gly Asp Ser 
                 
                     
                     
                 
                     
                   Arg Gly Ser Leu Leu Ser Pro Arg Pro Ile Ser Tyr 
                 
                     
                     
                 
                     
                   Leu Lys Gly Ser Ser Gly Gly Pro Leu Leu Cys Pro 
                 
                     
                     
                 
                     
                   Ala Gly His Ala Val Gly Ile Phe Arg Ala Ala Val 
                 
                     
                     
                 
                     
                   Cys Thr Arg Gly Val Ala Lys Ala Val Asp Phe Ile 
                 
                     
                     
                 
                     
                   Pro Val Glu Asn Leu Glu Thr Thr Met Arg. 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         11 . The fusion protein of  claim 6 , wherein said fusion partner is ubiquitin.  
     
     
         12 . A composition comprising a polynucleotide which encodes only the HCV protease or an active HCV protease analog.  
     
     
         13 . The composition of  claim 12 , wherein said polynucleotide encodes the HCV protease of FIG. 1.  
     
     
         14 . A composition comprising a polynucleotide which encodes a fusion protein comprising: 
 HCV protease or HCV protease analog; and    a fusion partner.    
     
     
         15 . The composition of  claim 14 , wherein said fusion partner is selected from the group consisting of hSOD, yeast a-factor, IL-2S, ubiquitin, β-galactosidase, β-lactamase, horseradish peroxidase, glucose oxidase, and urease.  
     
     
         16 . The composition of  claim 14 , wherein said HCV protease or HCV protease analog comprises a polypeptide having substantially the following sequence:  
       
         
           
                 
                 
               
                     
                     
                 
                     
                   Arg Arg Gly Arg Glu Ile Leu Leu Gly Pro Ala Asp 
                 
                     
                     
                 
                     
                   Gly Met Val Ser Lys Gly Trp Arg Leu Leu Ala Pro 
                 
                     
                     
                 
                     
                   Ile Thr Ala Tyr Ala Gln Gln Thr Arg Gly Leu Leu 
                 
                     
                     
                 
                     
                   Gly Cys Ile Ile Thr Ser Leu Thr Gly Arg Asp Lys 
                 
                     
                     
                 
                     
                   Asn Gln Val Glu Gly Glu Val Gln Ile Val Ser Thr 
                 
                     
                     
                 
                     
                   Ala Ala Gln Thr Phe Leu Ala Thr Cys Ile Asn Gly 
                 
                     
                     
                 
                     
                   Val Cys Trp Thr Val Tyr His Gly Ala Gly Thr Arg 
                 
                     
                     
                 
                     
                   Thr Ile Ala Ser Pro Lys Gly Pro Val Ile Gln Met 
                 
                     
                     
                 
                     
                   Tyr Thr Asn Val Asp Gln Asp Leu Val Gly Trp Pro 
                 
                     
                     
                 
                     
                   Ala Pro Gln Gly Ser Arg Ser Leu Thr Pro Cys Thr 
                 
                     
                     
                 
                     
                   Cys Gly Ser Ser Asp Leu Tyr Leu Val Thr Arg His 
                 
                     
                     
                 
                     
                   Ala Asp Val Ile Pro Val Arg Arg Arg Gly Asp Ser 
                 
                     
                     
                 
                     
                   Arg Gly Ser Leu Leu Ser Pro Arg Pro Ile Ser Tyr 
                 
                     
                     
                 
                     
                   Leu Lys Gly Ser Ser Gly Gly Pro Leu Leu Cys Pro 
                 
                     
                     
                 
                     
                   Ala Gly His Ala Val Gly Ile Phe Arg Ala Ala Val 
                 
                     
                     
                 
                     
                   Cys Thr Arg Gly Val Ala Lys Ala Val Asp Phe Ile 
                 
                     
                     
                 
                     
                   Pro Val Glu Asn Leu Glu Thr Thr Met Arg. 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         17 . The composition of  claim 14 , wherein said HCV protease or analog comprises a polypeptide having substantially the sequence:  
       
         
           
                 
                 
               
                     
                     
                 
                     
                   Gly Thr Tyr Val Tyr Asn His Leu Thr Pro Leu Arg 
                 
                     
                     
                 
                     
                   Asp Trp Ala His Asn Gly Leu Arg Asp Leu Ala Val 
                 
                     
                     
                 
                     
                   Ala Val Glu Pro Val Val Phe Ser Gln Met Glu Thr 
                 
                     
                     
                 
                     
                   Lys Leu Ile Thr Trp Gly Ala Asp Thr Ala Ala Cys 
                 
                     
                     
                 
                     
                   Gly Asp Ile Ile Asn Gly Leu Pro Val Ser Ala Arg 
                 
                     
                     
                 
                     
                   Arg Gly Arg Glu Ile Leu Leu Gly Pro Ala Asp Gly 
                 
                     
                     
                 
                     
                   Met Val Ser Lys Gly Trp Arg Leu Leu Ala Pro Ile 
                 
                     
                     
                 
                     
                   Thr Ala Tyr Ala Gln Gln Thr Arg Gly Leu Leu Gly 
                 
                     
                     
                 
                     
                   Cys Ile Ile Thr Ser Leu Thr Gly Arg Asp Lys Asn 
                 
                     
                     
                 
                     
                   Gln Val Glu Gly Glu Val Gln Ile Val Ser Thr Ala 
                 
                     
                     
                 
                     
                   Ala Gln Thr Phe Leu Ala Thr Cys Ile Ile Asn Gly 
                 
                     
                     
                 
                     
                   Val Cys Trp Thr Val Tyr His Gly Ala Gly Thr Arg 
                 
                     
                     
                 
                     
                   Thr Ile Ala Ser Pro Lys Gly Pro Val Ile Gln Met 
                 
                     
                     
                 
                     
                   Tyr Thr Asn Val Asp Gln Asp Leu Val Gly Trp Pro 
                 
                     
                     
                 
                     
                   Ala Ser Gln Gly Thr Arg Ser Leu Thr Pro Cys Thr 
                 
                     
                     
                 
                     
                   Cys Gly Ser Ser Asp Leu Tyr Leu Val Thr Arg His 
                 
                     
                     
                 
                     
                   Ala Asp Val Ile Pro Val Arg Arg Arg Gly Asp Ser 
                 
                     
                     
                 
                     
                   Arg Gly Ser Leu Leu Ser Pro Arg Pro Ile Ser Tyr 
                 
                     
                     
                 
                     
                   Leu Lys Gly Ser Ser Gly Gly Pro Leu Leu Cys Pro 
                 
                     
                     
                 
                     
                   Ala Gly His Ala Val Gly Ile Phe Arg Ala Ala Val 
                 
                     
                     
                 
                     
                   Cys Thr Arg Gly Val Ala Lys Ala Val Asp Phe Ile 
                 
                     
                     
                 
                     
                   Pro Val Glu Asn Leu Glu Thr Thr Met Arg Ser Pro 
                 
                     
                     
                 
                     
                   Val Phe Thr Asp Asn Ser Ser Pro Pro Val Val Pro 
                 
                     
                     
                 
                     
                   Gln Ser Phe Gln Val Ala His Leu His Ala Pro Thr 
                 
                     
                     
                 
                     
                   Gly Ser Gly Lys Ser Thr Lys Val Pro Ala Ala. 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         18 . The composition of  claim 14 , wherein said polypeptide has substantially the sequence:  
       
         
           
                 
                 
               
                     
                     
                 
                     
                   Gly Thr Tyr Val Tyr Asn His Leu Thr Pro Leu Arg 
                 
                     
                     
                 
                     
                   Asp Trp Ala His Asn Gly Leu Arg Asp Leu Ala Val 
                 
                     
                     
                 
                     
                   Ala Val Glu Pro Val Val Phe Ser Gln Met Glu Thr 
                 
                     
                     
                 
                     
                   Lys Leu Ile Thr Trp Gly Ala Asp Thr Ala Ala Cys 
                 
                     
                     
                 
                     
                   Gly Asp Ile Ile Asn Gly Leu Pro Val Ser Ala Arg 
                 
                     
                     
                 
                     
                   Arg Gly Arg Glu Ile Leu Leu Gly Pro Ala Asp Gly 
                 
                     
                     
                 
                     
                   Met Val Ser Lys Gly Trp Arg Leu Leu Ala Pro Ile 
                 
                     
                     
                 
                     
                   Thr Ala Tyr Ala Gln Gln Thr Arg Gly Leu Leu Gly 
                 
                     
                     
                 
                     
                   Cys Ile Ile Thr Ser Leu Thr Gly Arg Asp Lys Asn 
                 
                     
                     
                 
                     
                   Gln Val Glu Gly Glu Val Gln Ile Val Ser Thr Ala 
                 
                     
                     
                 
                     
                   Ala Gln Thr Phe Leu Ala Thr Cys Ile Ile Asn Gly 
                 
                     
                     
                 
                     
                   Val Cys Trp Thr Val Tyr His Gly Ala Gly Thr Arg 
                 
                     
                     
                 
                     
                   Thr Ile Ala Ser Pro Lys Gly Pro Val Ile Gln Met 
                 
                     
                     
                 
                     
                   Tyr Thr Asn Val Asp Gln Asp Leu Val Gly Trp Pro 
                 
                     
                     
                 
                     
                   Ala Ser Gln Gly Thr Arg Ser Leu Thr Pro Cys Thr 
                 
                     
                     
                 
                     
                   Cys Gly Ser Ser Asp Leu Tyr Leu Val Thr Arg His 
                 
                     
                     
                 
                     
                   Ala Asp Val Ile Pro Val Arg. 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         19 . A method for assaying compounds for activity against hepatitis C virus, comprising: 
 providing an active hepatitis C virus protease;    contacting said protease with a compound capable of inhibiting serine protease activity; and    measuring inhibition of the proteolytic activity of said hepatitis C virus protease.    
     
     
         20 . An expression vector for producing HCV protease or HCV protease analogs in a host cell, which vector comprises: 
 a polynucleotide encoding HCV protease or an HCV analog;    transcriptional and translational regulatory sequences functional in said host cell operably linked to said HCV protease-encoding polynucleotide; and    a selectable marker.    
     
     
         21 . The vector of  claim 20 , which further comprises a sequence encoding a fusion partner, linked to said HCV protease-encoding polynucleotide to form a fusion protein upon expression.  
     
     
         22 . The vector of  claim 21 , wherein said fusion partner is selected from the group consisting of hSOD, yeast α-factor, IL-2S, ubiquitin, β-galactosidase, β-lactamase, horseradish peroxidase, glucose oxidase, and urease.  
     
     
         23 . The vector of  claim 22 , where in said fusion partner is selected from the group consisting of ubiquitin, hSOD, and yeast α-factor.  
     
     
         24 . The vector of  claim 20 , wherein said HCV protease-encoding polynucleotide encodes a polypeptide having the substantially the following sequence:  
       
         
           
                 
                 
               
                     
                     
                 
                     
                   Arg Arg Gly Arg Glu Ile Leu Leu Gly Pro Ala Asp 
                 
                     
                     
                 
                     
                   Gly Met Val Ser Lys Gly Trp Arg Leu Leu Ala Pro 
                 
                     
                     
                 
                     
                   Ile Thr Ala Tyr Ala Gln Gln Thr Arg Gly Leu Leu 
                 
                     
                     
                 
                     
                   Gly Cys Ile Ile Thr Ser Leu Thr Gly Arg Asp Lys 
                 
                     
                     
                 
                     
                   Asn Gln Val Glu Gly Glu Val Gln Ile Val Ser Thr 
                 
                     
                     
                 
                     
                   Ala Ala Gln Thr Phe Leu Ala Thr Cys Ile Asn Gly 
                 
                     
                     
                 
                     
                   Val Cys Trp Thr Val Tyr His Gly Ala Gly Thr Arg 
                 
                     
                     
                 
                     
                   Thr Ile Ala Ser Pro Lys Gly Pro Val Ile Gln Met 
                 
                     
                     
                 
                     
                   Tyr Thr Asn Val Asp Gln Asp Leu Val Gly Trp Pro 
                 
                     
                     
                 
                     
                   Ala Pro Gln Gly Ser Arg Ser Leu Thr Pro Cys Thr 
                 
                     
                     
                 
                     
                   Cys Gly Ser Ser Asp Leu Tyr Leu Val Thr Arg His 
                 
                     
                     
                 
                     
                   Ala Asp Val Ile Pro Val Arg Arg Arg Gly Asp Ser 
                 
                     
                     
                 
                     
                   Arg Gly Ser Leu Leu Ser Pro Arg Pro Ile Ser Tyr 
                 
                     
                     
                 
                     
                   Leu Lys Gly Ser Ser Gly Gly Pro Leu Leu Cys Pro 
                 
                     
                     
                 
                     
                   Ala Gly His Ala Val Gly Ile Phe Arg Ala Ala Val 
                 
                     
                     
                 
                     
                   Cys Thr Arg Gly Val Ala Lys Ala Val Asp Phe Ile 
                 
                     
                     
                 
                     
                   Pro Val Glu Asn Leu Glu Thr Thr Met Arg. 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         25 . The vector of  claim 20 , wherein said HCV protease-encoding polynucleotide encodes a polypeptide having the substantially the following sequence:  
       
         
           
                 
                 
               
                     
                     
                 
                     
                   Gly Thr Tyr Val Tyr Asn His Leu Thr Pro Leu Arg 
                 
                     
                     
                 
                     
                   Asp Trp Ala His Asn Gly Leu Arg Asp Leu Ala Val 
                 
                     
                     
                 
                     
                   Ala Val Glu Pro Val Val Phe Ser Gln Met Glu Thr 
                 
                     
                     
                 
                     
                   Lys Leu Ile Thr Trp Gly Ala Asp Thr Ala Ala Cys 
                 
                     
                     
                 
                     
                   Gly Asp Ile Ile Asn Gly Leu Pro Val Ser Ala Arg 
                 
                     
                     
                 
                     
                   Arg Gly Arg Glu Ile Leu Leu Gly Pro Ala Asp Gly 
                 
                     
                     
                 
                     
                   Met Val Ser Lys Gly Trp Arg Leu Leu Ala Pro Ile 
                 
                     
                     
                 
                     
                   Thr Ala Tyr Ala Gln Gln Thr Arg Gly Leu Leu Gly 
                 
                     
                     
                 
                     
                   Cys Ile Ile Thr Ser Leu Thr Gly Arg Asp Lys Asn 
                 
                     
                     
                 
                     
                   Gln Val Glu Gly Glu Val Gln Ile Val Ser Thr Ala 
                 
                     
                     
                 
                     
                   Ala Gln Thr Phe Leu Ala Thr Cys Ile Ile Asn Gly 
                 
                     
                     
                 
                     
                   Val Cys Trp Thr Val Tyr His Gly Ala Gly Thr Arg 
                 
                     
                     
                 
                     
                   Thr Ile Ala Ser Pro Lys Gly Pro Val Ile Gln Met 
                 
                     
                     
                 
                     
                   Tyr Thr Asn Val Asp Gln Asp Leu Val Gly Trp Pro 
                 
                     
                     
                 
                     
                   Ala Ser Gln Gly Thr Arg Ser Leu Thr Pro Cys Thr 
                 
                     
                     
                 
                     
                   Cys Gly Ser Ser Asp Leu Tyr Leu Val Thr Arg His 
                 
                     
                     
                 
                     
                   Ala Asp Val Ile Pro Val Arg. 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         26 . The vector of  claim 20 , wherein said HCV protease-encoding polynucleotide encodes a polypeptide having the substantially the following sequence:  
       
         
           
                 
                 
               
                     
                     
                 
                     
                   Gly Thr Tyr Val Tyr Asn His Leu Thr Pro Leu Arg 
                 
                     
                     
                 
                     
                   Asp Trp Ala His Asn Gly Leu Arg Asp Leu Ala Val 
                 
                     
                     
                 
                     
                   Ala Val Glu Pro Val Val Phe Ser Gln Met Glu Thr 
                 
                     
                     
                 
                     
                   Lys Leu Ile Thr Trp Gly Ala Asp Thr Ala Ala Cys 
                 
                     
                     
                 
                     
                   Gly Asp Ile Ile Asn Gly Leu Pro Val Ser Ala Arg 
                 
                     
                     
                 
                     
                   Arg Gly Arg Glu Ile Leu Leu Gly Pro Ala Asp Gly 
                 
                     
                     
                 
                     
                   Met Val Ser Lys Gly Trp Arg Leu Leu Ala Pro Ile 
                 
                     
                     
                 
                     
                   Thr Ala Tyr Ala Gln Gln Thr Arg Gly Leu Leu Gly 
                 
                     
                     
                 
                     
                   Cys Ile Ile Thr Ser Leu Thr Gly Arg Asp Lys Asn 
                 
                     
                     
                 
                     
                   Gln Val Glu Gly Glu Val Gln Ile Val Ser Thr Ala 
                 
                     
                     
                 
                     
                   Ala Gln Thr Phe Leu Ala Thr Cys Ile Ile Asn Gly 
                 
                     
                     
                 
                     
                   Val Cys Trp Thr Val Tyr His Gly Ala Gly Thr Arg 
                 
                     
                     
                 
                     
                   Thr Ile Ala Ser Pro Lys Gly Pro Val Ile Gln Met 
                 
                     
                     
                 
                     
                   Tyr Thr Asn Val Asp Gln Asp Leu Val Gly Trp Pro 
                 
                     
                     
                 
                     
                   Ala Ser Gln Gly Thr Arg Ser Leu Thr Pro Cys Thr 
                 
                     
                     
                 
                     
                   Cys Gly Ser Ser Asp Leu Tyr Leu Val Thr Arg His 
                 
                     
                     
                 
                     
                   Ala Asp Val Ile Pro Val Arg Arg Arg Gly Asp Ser 
                 
                     
                     
                 
                     
                   Arg Gly Ser Leu Leu Ser Pro Arg Pro Ile Ser Tyr 
                 
                     
                     
                 
                     
                   Leu Lys Gly Ser Ser Gly Gly Pro Leu Leu Cys Pro 
                 
                     
                     
                 
                     
                   Ala Gly His Ala Val Gly Ile Phe Arg Ala Ala Val 
                 
                     
                     
                 
                     
                   Cys Thr Arg Gly Val Ala Lys Ala Val Asp Phe Ile 
                 
                     
                     
                 
                     
                   Pro Val Glu Asn Leu Glu Thr Thr Met Arg Ser Pro 
                 
                     
                     
                 
                     
                   Val Phe Thr Asp Asn Ser Ser Pro Pro Val Val Pro 
                 
                     
                     
                 
                     
                   Gln Ser Phe Gln Val Ala His Leu His Ala Pro Thr 
                 
                     
                     
                 
                     
                   Gly Ser Gly Lys Ser Thr Lys Val Pro Ala Ala.

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