US2013197191A1PendingUtilityA1

Binding Proteins That Bind to Human FGFR1c, Human Beta-Klotho and Both Human FGFR1c and Human Beta-Klotho

49
Assignee: AMGEN INCPriority: Dec 2, 2009Filed: Jan 10, 2013Published: Aug 1, 2013
Est. expiryDec 2, 2029(~3.4 yrs left)· nominal 20-yr term from priority
A61P 3/10A61P 9/10A61P 3/06A61P 3/00A61P 3/04C07K 2319/70A61P 1/18A61K 38/00C07K 14/705A61P 1/16C07K 2319/75C07K 14/001C07K 14/50
49
PatentIndex Score
0
Cited by
0
References
0
Claims

Abstract

Binding proteins that specifically bind to β-Klotho or portions thereof, FGFR1c or portions thereof, or both FGFR1c and β-Klotho, and optionally other proteins as well are provided. Coding sequences, methods of treatment and pharmaceutical compositions are also provided.

Claims

exact text as granted — not AI-modified
What is claimed is: 
     
         1 . A binding protein that selectively binds β-Klotho, wherein the binding protein comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 254) 
                 
                 
               
                     C [AGP][APS][DGN][EQ]F[QRT] C [GNRS][GNS][GT][GKQS][- 
                 
                     
                 
                   IK] C [ILV]P[LQRV][AEHP]W[LRV] C DG[DLV][DN]D C GD[GN] 
                 
                     
                 
                   SDE[AEKPS][GLPS][-AEV][-IT] C ; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 597) 
                 
                 
               
                     C LPDEFQ C [SG]SGR C IPQ[HT]W[VL] C DGLND C GDGSDE[PS]PA 
                 
                     
                 
                   [HT] C ; 
                 
                   and 
                 
                     
                 
                 
               
                   (SEQ ID NO: 249) 
                 
                 
               
                     C [AGPR][AP][DGNS][EQ]F[QRT] C [GSK]N[GT][GHR][R] C   
                 
                     
                 
                   [ILV][PS][ALQ][NST]W[LRV] C DG[DEV]DD C [GL]DGSDE[AET] 
                 
                     
                 
                   [DNS][-A][-T] C ; 
                 
             
                
               
            
             
                
                
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
               
            
           
         
         wherein amino acids in brackets indicate alternative amino acids at a specified position and “-” indicates no amino acid at the specified position. 
       
     
     
         2 . The binding protein of  claim 1 , wherein the binding protein comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 255) 
                 
                 
                 
               
                     
                   CGPDQFRCSSGKCIPQHWLCDGLNDCGDGSDESPATC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 256) 
                 
                 
                 
               
                     
                   CGANEFQCRSTGICVPVEWVCDGDNDCGDGSDEPPVC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 257) 
                 
                 
                 
               
                     
                   CGADQFRCGNGSCVPRAWRCDGVDDCGDGSDEAPEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 258) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 259) 
                 
                 
                 
               
                     
                   CASGEFTCNNGQCVPLAWRCDGVNDCQDGSDEKGC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 260) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 261) 
                 
                 
                 
               
                     
                   CPPDEFQCRGTKKCLPLAWVCDGDNDCEDDSDEESC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1237) 
                 
                 
                 
               
                     
                   CLPDEFQCGSGRCIPQHWLCDGLNDCGDGSDEPPAHC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1326) 
                 
                 
                 
               
                     
                   CLPDEFQCGSGRCIPQHWLCDGLNDCGDGSDEPPAHC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1327) 
                 
                 
                 
               
                     
                   CAPGEFTCKNTGRCIPLNWRCDGDDDCGDGSDETDC; 
                 
                     
                   and 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1328) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         3 . A binding protein that selectively binds β-Klotho, wherein the binding protein comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 394) 
                 
                 
               
                     C QSNEFR C RSGR C IPQHWL C DGLND C GDGSD 
                 
                     
                 
                   E[PS][PQ][AQ][-H]C; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 217) 
                 
                 
               
                     C [ELPQR][APS][DGIN][EGQ][FQ][FKPQRT][- 
                 
                     
                 
                   E] C [GNRS][NS]G[HNQR] C [IV]P[AELPQRV][AHPQRT]W 
                 
                     
                 
                   [LRV] C DG[DEV][DNP]D C [GLQ]D[DGNS]SDE[AEKT] 
                 
                     
                 
                   [DGLNS][-A][-H] C ; 
                 
                   and 
                 
                     
                 
                 
               
                   (SEQ ID NO: 254) 
                 
                 
               
                     C [AGP][APS][DGN][EQ]F[QRT] C [GNRS][GNS][GT][GKQS][- 
                 
                     
                 
                   IK] C [ILV]P[LQRV][AEHP]W[LRV] C DG[DLV][DN]D C GD 
                 
                     
                 
                   [GN]SDE[AEKPS][GLPS][-AEV][-IT] C ; 
                 
             
                
               
            
             
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
               
            
           
         
         wherein amino acids in brackets indicate alternative amino acids at a specified position and “-” indicates no amino acid at the specified position. 
       
     
     
         4 . The binding protein of  claim 3 , wherein the binding protein comprises one or more of: CQSNEFRCRSGRCIPQHWLCDGLNDCGDGSDESQQC (SEQ ID NO: 159);
 CRAGEFRCSNGRCVPLTWLCDGEDDCQDNSDEKNC (SEQ ID NO:1323);   CPSNQFPCRSTGICIPLAWVCDGLNDCGDGSDESPAHC (SEQ ID NO:1324); and   CQSNEFRCRSGRCIPQHWLCDGLNDCGDGSDEPPAHC (SEQ ID NO:1325).   
     
     
         5 . A binding protein that selectively binds FGFR1c, wherein the binding protein comprises: 
       
         
           
                 
               
                   (SEQ ID NO: 108) 
                 
                 
               
                     CG [AE][GS] LFTC [GR][NRS][AST][KN] ICIS [EHQS][AV]W 
                 
                     
                 
                   [IV] CDG [IV] DDC [DE] DNSDE [DKMNT][NSY] C ; 
                 
             
                
               
            
             
                
                
                
               
            
           
         
         wherein amino acids in brackets indicate alternative amino acids at a specified position and “-” indicates no amino acid at the specified position. 
       
     
     
         6 . The binding protein of  claim 5 , wherein the binding protein comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 109) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TNICISQVWV CDGVDDCEDN SDEDSC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 110) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TNICISQAWV CDGVDDCEDN SDENYC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 111) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TNICISQAWV CDGVDDCEDN SDETNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 112) 
                 
                 
                 
               
                     
                   CGEGLFTCGS TNICISSAWV CDGVDDCEDN SDENNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 113) 
                 
                 
                 
               
                     
                   CGEGLFTCRS TNICISHAWV CDGVDDCEDN SDENNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 114) 
                 
                 
                 
               
                     
                   CGEGLFTCRS TNICISEAWI CDGVDDCEDN SDEKNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 115) 
                 
                 
                 
               
                     
                   CGAGLFTCRS AKICISHAWV CDGIDDCEDN SDENNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 116) 
                 
                 
                 
               
                     
                   CGAGLFTCRN SKICISQAWV CDGVDDCDDN SDEKYC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 117) 
                 
                 
                 
               
                     
                   CGASLFTCRR SNICISQAWV CDGVDDCEDN SDEMNC; 
                 
                     
                   and 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 118) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TKICISQAWV CDGVDDCEDN SDEKNC. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         7 . The binding protein of  claim 6 , wherein the binding protein further comprises a linker comprising the sequence SAPASEPPGSL (SEQ ID NO: 12). 
     
     
         8 . The binding protein of  claim 7 , wherein the binding protein comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 293) 
                 
                 
               
                   CGAGLFTCRSTNICISQVWVCDGVDDCEDNSDEDSCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 294) 
                 
                 
               
                   CGAGLFTCRSTNICISQAWVCDGVDDCEDNSDENYCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 295) 
                 
                 
               
                   CGAGLFTCRSTNICISQAWVCDGVDDCEDNSDETNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 448) 
                 
                 
               
                   CGEGLFTCGSTNICISSAWVCDGVDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 297) 
                 
                 
               
                   CGEGLFTCRSTNICISHAWVCDGVDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 449) 
                 
                 
               
                   CGEGLFTCRSTNICISEAWICDGVDDCEDNSDEKNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 450) 
                 
                 
               
                   CGAGLFTCRSAKICISHAWVCDGIDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 464) 
                 
                 
               
                   CGAGLFTCRNSKICISQAWVCDGVDDCDDNSDEKYCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 296) 
                 
                 
               
                   CGASLFTCRRSNICISQAWVCDGVDDCEDNSDEMNCSAPASEPPGSL; 
                 
                   and 
                 
                     
                 
                 
               
                   (SEQ ID NO: 287) 
                 
                 
               
                   CGAGLFTCRSTKICISQAWVCDGVDDCEDNSDEKNCSAPASEPPGSL. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         9 . The binding protein of  claim 8 , wherein the binding protein comprises the sequence: 
       
         
           
                 
               
                   (SEQ ID NO: 297) 
                 
                 
               
                   CGEGLFTCRSTNICISHAWVCDGVDDCEDNSDENNCSAPASEPPGSL. 
                 
             
                
               
            
             
                
               
            
           
         
       
     
     
         10 . A binding protein that selectively binds β-Klotho, wherein the binding protein comprises:
 (a) a first β-Klotho binding domain comprising a sequence selected from the group represented by the sequences: 
 
       
         
           
                 
               
                   (SEQ ID NO: 254) 
                 
                 
               
                     C [AGP][APS][DGN][EQ]F[QRT] C [GNRS][GNS][GT][GKQS][- 
                 
                     
                 
                   IK] C [ILV]P[LQRV][AEHP]W[LRV] C DG[DLV][DN]D C GD 
                 
                     
                 
                   [GN]SDE[AEKPS][GLPS][-AEV][-IT] C ; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 597) 
                 
                 
               
                     C LPDEFQ C [SG]SGR C IPQ[HT]W[VL] C DGLND C GDGSDE[PS]PA 
                 
                     
                 
                   [HT] C ; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 249) 
                 
                 
               
                     C [AGPR][AP][DGNS][EQ]F[QRT] C [GSK]N[GT][GHR][- 
                 
                     
                 
                   R] C [ILV][PS][ALQ][NST]W[LRV] C DG[DEV]DD C [GL]DGSDE 
                 
                     
                 
                   [AET][DNS][-A][-T] C ; 
                 
                   and 
                 
             
                
               
            
             
                
                
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
                
               
            
           
         
         (b) a second β-Klotho binding domain comprising a sequence selected from the group represented by the sequences: 
       
       
         
           
                 
               
                   (SEQ ID NO: 394) 
                 
                 
               
                     C QSNEFR C RSGR C IPQHWL C DGLND C GDGSD 
                 
                     
                 
                   E[PS][PQ][AQ][-H]C; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 217) 
                 
                 
               
                     C [ELPQR][APS][DGIN][EGQ][FQ][FKPQRT][- 
                 
                     
                 
                   E] C [GNRS][NS]G[HNQR] C [IV]P[AELPQRV][AHPQRT]W 
                 
                     
                 
                   [LRV] C DG[DEV][DNP]D C [GLQ]D[DGNS]SDE[AEKT] 
                 
                     
                 
                   [DGLNS][-A][-H] C ; 
                 
                   and 
                 
                     
                 
                 
               
                   (SEQ ID NO: 254) 
                 
                 
               
                     C [AGP][APS][DGN][EQ]F[QRT] C [GNRS][GNS][GT][GKQS][- 
                 
                     
                 
                   IK] C [ILV]P[LQRV][AEHP]W[LRV] C DG[DLV][DN]D C GD[GN] 
                 
                     
                 
                   SDE[AEKPS][GLPS][-AEV][-IT] C ; 
                 
             
                
               
            
             
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
               
            
           
         
         wherein amino acids in brackets indicate alternative amino acids at a specified position and “-” indicates no amino acid at the specified position. 
       
     
     
         11 . The binding protein of  claim 10  further comprising at least one linker joining (a) and (b). 
     
     
         12 . The binding protein of  claim 11 , wherein the linker is one or more of: ETPT (SEQ ID NO: 1319), GDSHILPFSTPGPST (SEQ ID NO: 14), SAPASEPPGSL (SEQ ID NO:12), PAPT (SEQ ID NO:1320), QAPT (SEQ ID NO:1321), AQPT (SEQ ID NO:1322), AHT, PERT (SEQ ID NO: 7), TTRT (SEQ ID NO: 8), GTTGPT (SEQ ID NO: 9), ETSGPT (SEQ ID NO: 10), SQDPEFHKVS (SEQ ID NO: 11), and GRPGPGATSAPAA (SEQ ID NO: 13). 
     
     
         13 . The binding protein of  claim 10 , wherein the β-Klotho binding domain of (a) comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 255) 
                 
                 
                 
               
                     
                   CGPDQFRCSSGKCIPQHWLCDGLNDCGDGSDESPATC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 256) 
                 
                 
                 
               
                     
                   CGANEFQCRSTGICVPVEWVCDGDNDCGDGSDEPPVC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 257) 
                 
                 
                 
               
                     
                   CGADQFRCGNGSCVPRAWRCDGVDDCGDGSDEAPEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 258) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 259) 
                 
                 
                 
               
                     
                   CASGEFTCNNGQCVPLAWRCDGVNDCQDGSDEKGC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 260) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 261) 
                 
                 
                 
               
                     
                   CPPDEFQCRGTKKCLPLAWVCDGDNDCEDDSDEESC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1237) 
                 
                 
                 
               
                     
                   CLPDEFQCGSGRCIPQHWLCDGLNDCGDGSDEPPAHC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1326) 
                 
                 
                 
               
                     
                   CLPDEFQCGSGRCIPQHWLCDGLNDCGDGSDEPPAHC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1327) 
                 
                 
                 
               
                     
                   CAPGEFTCKNTGRCIPLNWRCDGDDDCGDGSDETDC; 
                 
                     
                   and 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1328) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         14 . The binding protein of  claim 10 , wherein the β-Klotho binding domain of (b) comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 159) 
                 
                 
                 
               
                     
                   CQSNEFRCRSGRCIPQHWLCDGLNDCGDGSDESQQC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1323) 
                 
                 
                 
               
                     
                   CRAGEFRCSNGRCVPLTWLCDGEDDCQDNSDEKNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1324) 
                 
                 
                 
               
                     
                   CPSNQFPCRSTGICIPLAWVCDGLNDCGDGSDESPAHC; 
                 
                     
                   and 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1325) 
                 
                 
                 
               
                     
                   CQSNEFRCRSGRCIPQHWLCDGLNDCGDGSDEPPAHC. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         15 . A binding protein that selectively binds β-Klotho and FGFR1c comprising (a) one or more domains that selectively bind β-Klotho, and (b) one or more domains that selectively bind FGFR1c, wherein the binding protein comprises:
 (a) a β-Klotho binding domain comprising one or more of: 
 
       
         
           
                 
               
                   (SEQ ID NO: 254) 
                 
                 
               
                     C [AGP][APS][DGN][EQ]F[QRT] C [GNRS][GNS][GT][GKQS][- 
                 
                     
                 
                   IK] C [ILV]P[LQRV][AEHP]W[LRV] C DG[DLV][DN]D C GD[GN]S 
                 
                     
                 
                   DE[AEKPS][GLPS][-AEV][-IT] C ; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 597) 
                 
                 
               
                     C LPDEFQ C [SG]SGR C IPQ[HT]W[VL] C DGLND C GDGSDE[PS]PA 
                 
                     
                 
                   [HT] C ; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 249) 
                 
                 
               
                     C [AGPR][AP][DGNS][EQ]F[QRT] C [GSK]N[GT][GHR][- 
                 
                     
                 
                   R] C [ILV][PS][ALQ][NST]W[LRV] C DG[DEV]DD C [GL]DGSDE 
                 
                     
                 
                   [AET][DNS][-A][-T] C ; 
                 
             
                
               
            
             
                
                
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
               
            
           
         
         (b) an FGFR1c binding domain comprising one or more of: 
       
       
         
           
                 
               
                   (SEQ ID NO: 108) 
                 
                 
               
                     CG [AE][GS] LFTC [GR][NRS][AST][KN] ICIS [EHQS][AV]W 
                 
                     
                 
                   [IV] CDG [IV] DDC [DE] DNSDE [DKMNT][NSY] C ; 
                 
                   and 
                 
             
                
               
            
             
                
                
                
                
               
            
           
         
         wherein amino acids in brackets indicate alternative amino acids at a specified position and “-” indicates no amino acid at the specified position. 
       
     
     
         16 . The binding protein of  claim 10  further comprising at least one linker joining (a) and (b). 
     
     
         17 . The binding protein of  claim 16 , wherein the linker is one or more of: ETPT (SEQ ID NO: 1319), GDSHILPFSTPGPST (SEQ ID NO: 14), SAPASEPPGSL (SEQ ID NO:12), PAPT (SEQ ID NO:1320), QAPT (SEQ ID NO:1321), AQPT (SEQ ID NO:1322), AHT, PERT (SEQ ID NO: 7), TTRT (SEQ ID NO: 8), GTTGPT (SEQ ID NO: 9), ETSGPT (SEQ ID NO: 10), SQDPEFHKVS (SEQ ID NO: 11), and GRPGPGATSAPAA (SEQ ID NO: 13). 
     
     
         18 . The binding protein of  claim 15 , wherein
 (i) the β-Klotho binding domain of (a) comprises one or more of:   
       
         
           
                 
               
                   (SEQ ID NO: 255) 
                 
                 
                 
               
                     
                   CGPDQFRCSSGKCIPQHWLCDGLNDCGDGSDESPATC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 256) 
                 
                 
                 
               
                     
                   CGANEFQCRSTGICVPVEWVCDGDNDCGDGSDEPPVC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 257) 
                 
                 
                 
               
                     
                   CGADQFRCGNGSCVPRAWRCDGVDDCGDGSDEAPEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 258) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 259) 
                 
                 
                 
               
                     
                   CASGEFTCNNGQCVPLAWRCDGVNDCQDGSDEKGC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 260) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 261) 
                 
                 
                 
               
                     
                   CPPDEFQCRGTKKCLPLAWVCDGDNDCEDDSDEESC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1237) 
                 
                 
                 
               
                     
                   CLPDEFQCGSGRCIPQHWLCDGLNDCGDGSDEPPAHC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1326) 
                 
                 
                 
               
                     
                   CLPDEFQCGSGRCIPQHWLCDGLNDCGDGSDEPPAHC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1327) 
                 
                 
                 
               
                     
                   CAPGEFTCKNTGRCIPLNWRCDGDDDCGDGSDETDC; 
                 
                     
                   and 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1328) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC; 
                 
                     
                   and 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
               
            
           
         
       
     
     
         19 . The binding protein of  claim 18 , wherein the FGFR1c binding domain of (b) further comprises a linker comprising the sequence SAPASEPPGSL (SEQ ID NO: 12). 
     
     
         20 . The binding protein of  claim 19 , wherein the FGFR1c binding domain comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 293) 
                 
                 
               
                   CGAGLFTCRSTNICISQVWVCDGVDDCEDNSDEDSCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 294) 
                 
                 
               
                   CGAGLFTCRSTNICISQAWVCDGVDDCEDNSDENYCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 295) 
                 
                 
               
                   CGAGLFTCRSTNICISQAWVCDGVDDCEDNSDETNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 448) 
                 
                 
               
                   CGEGLFTCGSTNICISSAWVCDGVDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 297) 
                 
                 
               
                   CGEGLFTCRSTNICISHAWVCDGVDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 449) 
                 
                 
               
                   CGEGLFTCRSTNICISEAWICDGVDDCEDNSDEKNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 450) 
                 
                 
               
                   CGAGLFTCRSAKICISHAWVCDGIDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 464) 
                 
                 
               
                   CGAGLFTCRNSKICISQAWVCDGVDDCDDNSDEKYCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 296) 
                 
                 
               
                   CGASLFTCRRSNICISQAWVCDGVDDCEDNSDEMNCSAPASEPPGSL; 
                 
                   and 
                 
                     
                 
                 
               
                   (SEQ ID NO: 287) 
                 
                 
               
                   CGAGLFTCRSTKICISQAWVCDGVDDCEDNSDEKNCSAPASEPPGSL. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         21 . A binding protein that selectively binds β-Klotho and FGFR1c comprising (a) one or more domains that selectively bind β-Klotho, and (b) one or more domains that selectively bind FGFR1c, wherein the binding protein comprises:
 (a) a β-Klotho binding domain comprising one or more of: 
 
       
         
           
                 
               
                   (SEQ ID NO: 394) 
                 
                 
               
                     C QSNEFR C RSGR C IPQHWL C DGLND C GDGSD 
                 
                     
                 
                   E[PS][PQ][AQ][-H]C; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 217) 
                 
                 
               
                     C [ELPQR][APS][DGIN][EGQ][FQ][FKPQRT][- 
                 
                     
                 
                   E] C [GNRS][NS]G[HNQR] C [IV]P[AELPQRV][AHPQRT]W[LRV] C   
                 
                     
                 
                   DG[DEV][DNP]D C [GLQ]D[DGNS]SDE[AEKT][DGLNS][-A] 
                 
                     
                 
                   [-H] C ; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 254) 
                 
                 
               
                     C [AGP][APS][DGN][EQ]F[QRT] C [GNRS][GNS][GT][GKQS][- 
                 
                     
                 
                   IK] C [ILV]P[LQRV][AEHP]W[LRV] C DG[DLV][DN]D C GD[GN]SD 
                 
                     
                 
                   E[AEKPS][GLPS][-AEV][-IT] C ; 
                 
             
                
               
            
             
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
               
            
           
         
         
           and 
         
         (b) an FGFR1c binding domain comprising a sequence selected from the group represented by the sequence: 
       
       
         
           
                 
               
                   (SEQ ID NO: 108) 
                 
                 
               
                     CG [AE][GS] LFTC [GR][NRS][AST][KN] ICIS [EHQS][AV]W 
                 
                     
                 
                   [IV] CDG [IV] DDC [DE] DNSDE [DKMNT][NSY] C ; 
                 
             
                
               
            
             
                
                
                
               
            
           
         
         wherein amino acids in brackets indicate alternative amino acids at a specified position and “-” indicates no amino acid at the specified position. 
       
     
     
         22 . The binding protein of  claim 10  further comprising at least one linker joining (a) and (b). 
     
     
         23 . The binding protein of  claim 22 , wherein the linker is one or more of: ETPT (SEQ ID NO: 1319), GDSHILPFSTPGPST (SEQ ID NO: 14), SAPASEPPGSL (SEQ ID NO:12), PAPT (SEQ ID NO:1320), QAPT (SEQ ID NO:1321), AQPT (SEQ ID NO:1322), AHT, PERT (SEQ ID NO: 7), TTRT (SEQ ID NO: 8), GTTGPT (SEQ ID NO: 9), ETSGPT (SEQ ID NO: 10), SQDPEFHKVS (SEQ ID NO: 11), and GRPGPGATSAPAA (SEQ ID NO: 13). 
     
     
         24 . The binding protein of  claim 21 , wherein
 (i) the β-Klotho binding domain of (a) comprises one or more of:   
       
         
           
                 
               
                   (SEQ ID NO: 159) 
                 
                 
                 
               
                     
                   CQSNEFRCRSGRCIPQHWLCDGLNDCGDGSDESQQC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1323) 
                 
                 
                 
               
                     
                   CRAGEFRCSNGRCVPLTWLCDGEDDCQDNSDEKNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1324) 
                 
                 
                 
               
                     
                   CPSNQFPCRSTGICIPLAWVCDGLNDCGDGSDESPAHC; 
                 
                     
                   and 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1325) 
                 
                 
                 
               
                     
                   CQSNEFRCRSGRCIPQHWLCDGLNDCGDGSDEPPAHC; 
                 
                     
                   and 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
               
            
           
         
         (ii) the FGFR1c binding domain of (b) comprises one or more of: 
       
       
         
           
                 
               
                   (SEQ ID NO: 109) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TNICISQVWV CDGVDDCEDN SDEDSC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 110) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TNICISQAWV CDGVDDCEDN SDENYC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 111) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TNICISQAWV CDGVDDCEDN SDETNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 112) 
                 
                 
                 
               
                     
                   CGEGLFTCGS TNICISSAWV CDGVDDCEDN SDENNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 113) 
                 
                 
                 
               
                     
                   CGEGLFTCRS TNICISHAWV CDGVDDCEDN SDENNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 114) 
                 
                 
                 
               
                     
                   CGEGLFTCRS TNICISEAWI CDGVDDCEDN SDEKNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 115) 
                 
                 
                 
               
                     
                   CGAGLFTCRS AKICISHAWV CDGIDDCEDN SDENNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 116) 
                 
                 
                 
               
                     
                   CGAGLFTCRN SKICISQAWV CDGVDDCDDN SDEKYC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 117) 
                 
                 
                 
               
                     
                   CGASLFTCRR SNICISQAWV CDGVDDCEDN SDEMNC; 
                 
                     
                   and 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 118) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TKICISQAWV CDGVDDCEDN SDEKNC. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         25 . The binding protein of  claim 24 , wherein the FGFR1c binding domain of (b) further comprises a linker comprising the sequence SAPASEPPGSL (SEQ ID NO: 12). 
     
     
         26 . The binding protein of  claim 25 , wherein the FGFR1c binding domain comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 293) 
                 
                 
               
                   CGAGLFTCRSTNICISQVWVCDGVDDCEDNSDEDSCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 294) 
                 
                 
               
                   CGAGLFTCRSTNICISQAWVCDGVDDCEDNSDENYCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 295) 
                 
                 
               
                   CGAGLFTCRSTNICISQAWVCDGVDDCEDNSDETNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 448) 
                 
                 
               
                   CGEGLFTCGSTNICISSAWVCDGVDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 297) 
                 
                 
               
                   CGEGLFTCRSTNICISHAWVCDGVDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 449) 
                 
                 
               
                   CGEGLFTCRSTNICISEAWICDGVDDCEDNSDEKNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 450) 
                 
                 
               
                   CGAGLFTCRSAKICISHAWVCDGIDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 464) 
                 
                 
               
                   CGAGLFTCRNSKICISQAWVCDGVDDCDDNSDEKYCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 296) 
                 
                 
               
                   CGASLFTCRRSNICISQAWVCDGVDDCEDNSDEMNCSAPASEPPGSL; 
                 
                   and 
                 
                     
                 
                 
               
                   (SEQ ID NO: 287) 
                 
                 
               
                   CGAGLFTCRSTKICISQAWVCDGVDDCEDNSDEKNCSAPASEPPGSL. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         27 . A binding protein that selectively binds β-Klotho and FGFR1c comprising (a) one or more domains that selectively bind β-Klotho, and (b) one or more domains that selectively bind FGFR1c, wherein the binding protein comprises:
 (a) a β-Klotho binding domain comprising:
 (i) a first binding domain comprising one or more of: 
 
 
       
         
           
                 
               
                   (SEQ ID NO: 254) 
                 
                 
               
                     C [AGP][APS][DGN][EQ]F[QRT] C [GNRS][GNS][GT][GKQS][- 
                 
                     
                 
                   IK] C [ILV]P[LQRV][AEHP]W[LRV] C DG[DLV][DN]D C GD[GN]SD 
                 
                     
                 
                   E[AEKPS][GLPS][-AEV][-IT] C ; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 597) 
                 
                 
               
                     C LPDEFQ C [SG]SGR C IPQ[HT]W[VL] C DGLND C GDGSDE[PS]PA 
                 
                     
                 
                   [HT] C ; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 249) 
                 
                 
               
                     C [AGPR][AP][DGNS][EQ]F[QRT] C [GSK]N[GT][GHR][- 
                 
                     
                 
                   R] C [ILV][PS][ALQ][NST]W[LRV] C DG[DEV]DD C [GL]DGSDE[A 
                 
                     
                 
                   ET][DNS][-A][-T] C ; 
                 
             
                
               
            
             
                
                
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
               
            
           
         
         
           (ii) a second binding domain comprising one or more of: 
         
       
       
         
           
                 
               
                   (SEQ ID NO: 394) 
                 
                 
               
                     C QSNEFR C RSGR C IPQHWL C DGLND C GDGSD 
                 
                     
                 
                   E[PS][PQ][AQ][-H]C; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 217) 
                 
                 
               
                     C [ELPQR][APS][DGIN][EGQ][FQ][FKPQRT][-E] C [GNRS] 
                 
                     
                 
                   [NS]G[HNQR] C [IV]P[AELPQRV][AHPQRT]W[LRV] C DG 
                 
                     
                 
                   [DEV][DNP]D C [GLQ]D[DGNS]SDE[AEKT][DGLNS][-A][-H] C ; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 254) 
                 
                 
               
                     C [AGP][APS][DGN][EQ]F[QRT] C [GNRS][GNS][GT][GKQS][- 
                 
                     
                 
                   IK] C [ILV]P[LQRV][AEHP]W[LRV] C DG[DLV][DN]D C GD[GN]SD 
                 
                     
                 
                   E[AEKPS][GLPS][-AEV][-IT] C ; 
                 
             
                
               
            
             
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
                
               
            
             
                
               
            
             
                
                
                
                
                
               
            
           
         
         
           
             and 
           
           (iii) a linker joining the first binding domain and the second binding domain; and 
         
         (b) an FGFR1c binding domain comprising a sequence selected from the group represented by the sequence: 
       
       
         
           
                 
               
                   (SEQ ID NO: 108) 
                 
                 
                 
               
                     
                     CG [AE][GS] LFTC [GR][NRS][AST][KN] ICIS [EHQS][AV]W 
                 
                     
                     
                 
                     
                   [IV] CDG [IV] DDC [DE] DNSDE [DKMNT][NSY] C ; 
                 
                     
                   and 
                 
             
                
               
            
             
                
                
                
                
               
            
           
         
         (c) a linker joining the sequences of (a) and (b), 
         wherein amino acids in brackets indicate alternative amino acids at a specified position and “-” indicates no amino acid at the specified position. 
       
     
     
         28 . The binding protein of  claim 27 , wherein the first binding domain of (a) comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 255) 
                 
                 
                 
               
                     
                   CGPDQFRCSSGKCIPQHWLCDGLNDCGDGSDESPATC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 256) 
                 
                 
                 
               
                     
                   CGANEFQCRSTGICVPVEWVCDGDNDCGDGSDEPPVC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 257) 
                 
                 
                 
               
                     
                   CGADQFRCGNGSCVPRAWRCDGVDDCGDGSDEAPEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 258) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 259) 
                 
                 
                 
               
                     
                   CASGEFTCNNGQCVPLAWRCDGVNDCQDGSDEKGC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 260) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 261) 
                 
                 
                 
               
                     
                   CPPDEFQCRGTKKCLPLAWVCDGDNDCEDDSDEESC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1237) 
                 
                 
                 
               
                     
                   CLPDEFQCGSGRCIPQHWLCDGLNDCGDGSDEPPAHC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1326) 
                 
                 
                 
               
                     
                   CLPDEFQCGSGRCIPQHWLCDGLNDCGDGSDEPPAHC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1327) 
                 
                 
                 
               
                     
                   CAPGEFTCKNTGRCIPLNWRCDGDDDCGDGSDETDC; 
                 
                     
                   and 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1328) 
                 
                 
                 
               
                     
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEIC. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         29 . The binding protein of  claim 27 , wherein the second binding domain of (a) comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 159) 
                 
                 
                 
               
                     
                   CQSNEFRCRSGRCIPQHWLCDGLNDCGDGSDESQQC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1323) 
                 
                 
                 
               
                     
                   CRAGEFRCSNGRCVPLTWLCDGEDDCQDNSDEKNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1324) 
                 
                 
                 
               
                     
                   CPSNQFPCRSTGICIPLAWVCDGLNDCGDGSDESPAHC; 
                 
                     
                   and 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 1325) 
                 
                 
                 
               
                     
                   CQSNEFRCRSGRCIPQHWLCDGLNDCGDGSDEPPAHC. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         30 . The binding protein of  claim 27 , wherein the linker of (a) comprises one or more of: ETPT (SEQ ID NO: 1319), GDSHILPFSTPGPST (SEQ ID NO: 14), SAPASEPPGSL (SEQ ID NO:12), PAPT (SEQ ID NO:1320) and QAPT (SEQ ID NO:1321). 
     
     
         31 . The binding protein of  claim 27 , wherein the FGFR1c binding domain comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 109) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TNICISQVWV CDGVDDCEDN SDEDSC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 110) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TNICISQAWV CDGVDDCEDN SDENYC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 111) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TNICISQAWV CDGVDDCEDN SDETNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 112) 
                 
                 
                 
               
                     
                   CGEGLFTCGS TNICISSAWV CDGVDDCEDN SDENNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 113) 
                 
                 
                 
               
                     
                   CGEGLFTCRS TNICISHAWV CDGVDDCEDN SDENNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 114) 
                 
                 
                 
               
                     
                   CGEGLFTCRS TNICISEAWI CDGVDDCEDN SDEKNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 115) 
                 
                 
                 
               
                     
                   CGAGLFTCRS AKICISHAWV CDGIDDCEDN SDENNC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 116) 
                 
                 
                 
               
                     
                   CGAGLFTCRN SKICISQAWV CDGVDDCDDN SDEKYC; 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 117) 
                 
                 
                 
               
                     
                   CGASLFTCRR SNICISQAWV CDGVDDCEDN SDEMNC; 
                 
                     
                   and 
                 
                     
                     
                 
                 
               
                   (SEQ ID NO: 118) 
                 
                 
                 
               
                     
                   CGAGLFTCRS TKICISQAWV CDGVDDCEDN SDEKNC. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         32 . The binding protein of  claim 31 , further comprising a linker comprising the sequence SAPASEPPGSL (SEQ ID NO: 12). 
     
     
         33 . The binding protein of  claim 32 , wherein the binding protein comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 293) 
                 
                 
               
                   CGAGLFTCRSTNICISQVWVCDGVDDCEDNSDEDSCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 294) 
                 
                 
               
                   CGAGLFTCRSTNICISQAWVCDGVDDCEDNSDENYCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 295) 
                 
                 
               
                   CGAGLFTCRSTNICISQAWVCDGVDDCEDNSDETNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 448) 
                 
                 
               
                   CGEGLFTCGSTNICISSAWVCDGVDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 297) 
                 
                 
               
                   CGEGLFTCRSTNICISHAWVCDGVDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 449) 
                 
                 
               
                   CGEGLFTCRSTNICISEAWICDGVDDCEDNSDEKNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 450) 
                 
                 
               
                   CGAGLFTCRSAKICISHAWVCDGIDDCEDNSDENNCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 464) 
                 
                 
               
                   CGAGLFTCRNSKICISQAWVCDGVDDCDDNSDEKYCSAPASEPPGSL; 
                 
                     
                 
                 
               
                   (SEQ ID NO: 296) 
                 
                 
               
                   CGASLFTCRRSNICISQAWVCDGVDDCEDNSDEMNCSAPASEPPGSL; 
                 
                   and 
                 
                     
                 
                 
               
                   (SEQ ID NO: 287) 
                 
                 
               
                   CGAGLFTCRSTKICISQAWVCDGVDDCEDNSDEKNCSAPASEPPGSL. 
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         34 . The binding protein of  claim 33 , wherein the FGFR1c binding domain comprises one or more of: 
       
         
           
                 
               
                   (SEQ ID NO: 297) 
                 
                 
               
                   CGEGLFTCRSTNICISHAWVCDGVDDCEDNSDENNCSAPASEPPGSL. 
                 
             
                
               
            
             
                
               
            
           
         
       
     
     
         35 . The binding protein of  claim 27 , wherein the linker of (c) comprises one or more of: AQPT (SEQ ID NO:1322), AHT, PERT (SEQ ID NO: 7), TTRT (SEQ ID NO: 8), GTTGPT (SEQ ID NO: 9), ETSGPT (SEQ ID NO: 10), SQDPEFHKVS (SEQ ID NO: 11), SAPASEPPGSL (SEQ ID NO: 12), GRPGPGATSAPAA (SEQ ID NO: 13), and GDSHILPFSTPGPST (SEQ ID NO: 14). 
     
     
         36 . The binding protein of  claim 27 , wherein the binding protein comprises the sequence: 
       
         
           
                 
               
                   (SEQ ID NO: 359) 
                 
                 
               
                   CGADQFRCGNGSCVPRAWRCDGVDDCGDGSDEAPEICETPTCQSNEFRCR 
                 
                     
                 
                   SGRCIPQHWLCDGLNDCGDGSDESQQCSAPASEPPGSLCGEGLFTCRSTN 
                 
                     
                 
                   ICISHAWVCDGVDDCEDNSDENNCSAPASEPPGSL. 
                 
             
                
               
            
             
                
                
                
                
                
               
            
           
         
       
     
     
         37 . The binding protein of  claim 27 , wherein the binding protein comprises the sequence: 
       
         
           
                 
               
                   (SEQ ID NO: 339) 
                 
                 
               
                   CLPDEFQCGSGRCIPQHWLCDGLNDCGDGSDEPPAHCSAPASEPPGSLCR 
                 
                     
                 
                   AGEFRCSNGRCVPLTWLCDGEDDCQDNSDEKNCAQPTCGEGLFTCRSTNI 
                 
                     
                 
                   CISHAWVCDGVDDCEDNSDENNCSAPASEPPGSL. 
                 
             
                
               
            
             
                
                
                
                
                
               
            
           
         
       
     
     
         38 . The binding protein of  claim 27 , wherein the binding protein comprises the sequence: 
       
         
           
                 
               
                   (SEQ ID NO: 1270) 
                 
                 
               
                   CAPGEFTCKNTGRCIPLNWRCDGDDDCGDGSDETDCPAPTCPSNQFPCRS 
                 
                     
                 
                   TGICIPLAWVCDGLNDCGDGSDESPAHCSAPASEPPGSLCGEGLFTCRST 
                 
                     
                 
                   NICISHAWVCDGVDDCEDNSDENNCSAPASEPPGSL. 
                 
             
                
               
            
             
                
                
                
                
                
               
            
           
         
       
     
     
         39 . The binding protein of  claim 27 , wherein the binding protein comprises the sequence: 
       
         
           
                 
               
                   (SEQ ID NO: 1271) 
                 
                 
               
                   CGPDEFRCNNGQCIPLPWRCDGVDDCGDNSDEPLEICQAPTCQSNEFRCR 
                 
                     
                 
                   SGRCIPQHWLCDGLNDCGDGSDEPPAHCSAPASEPPGSLCGEGLFTCRST 
                 
                     
                 
                   NICISHAWVCDGVDDCEDNSDENNCSAPASEPPGSL. 
                 
             
                
               
            
             
                
                
                
                
                
               
            
           
         
       
     
     
         40 . A binding protein comprising:
   (B1) a -(L1) x -(B2) b -(L2) y -(1C) c -(L3) z      wherein a, b, c, x, y, and z is 0, 1, or 2, provided however that at least one of a, b, and c is 1 or 2;   wherein B1 is an amino acid sequence as set forth in  claim 1 ;   wherein B2 is an amino acid sequence as set forth in  claim 3 ;   wherein 1C is an amino acid sequence as set forth in  claim 5 ;   wherein L1 is an amino acid sequence as set forth in  claim 12 ;   wherein L2 is an amino acid sequence as set forth in  claim 17 ; and   wherein L3 is SAPASEPPGSL.   
     
     
         41 . The binding protein of  claim 1 ,  3 ,  5 ,  10 ,  15 ,  21 ,  27 ,  36 ,  37 ,  38 , or  39 , wherein the binding protein further comprises a half life-extending moiety that increases the serum half-life of the binding protein in a mammal compared to the serum half-life of the binding protein lacking the half life-extending moiety in a mammal. 
     
     
         42 . The binding protein of  claim 41 , wherein the half life-extending moiety is one or more of: polyethylene glycol (PEG), human serum albumin (HSA), an immunoglobulin (IgG), and an Fc moeity. 
     
     
         43 . The binding protein of  claim 41 , wherein the half life-extending moiety comprises a protein sequence that specifically binds a blood factor. 
     
     
         44 . The binding protein of  claim 43 , wherein the blood factor is serum albumin, an immunoglobulin or an erythrocyte. 
     
     
         45 . The binding protein of  claim 1 ,  3 ,  5 ,  10 ,  15 ,  21 ,  27 ,  36 ,  37 ,  38 , or  39 , wherein the binding protein comprises an amino acid sequence comprising one or more non-naturally occurring amino acids. 
     
     
         46 . A polynucleotide encoding a binding protein comprising one or more of:
 (a) a β-Klotho binding domain;   (b) an FGFR1c binding domain;   (c) one or more β-Klotho binding domains and an FGFR1c binding domain; and   (c) a binding protein of  claim 1 ,  3 ,  5 ,  10 ,  15 ,  21 ,  27 ,  36 ,  37 ,  38 , or  39 .   
     
     
         47 . An expression vector comprising the polynucleotide of  claim 46 . 
     
     
         48 . A host cell comprising the expression vector of  claim 47 . 
     
     
         49 . A method of making a binding protein comprising incubating the host cell of  claim 48  under conditions suitable to induce expression of the binding protein, and optionally isolating the expressed polypeptide. 
     
     
         50 . A pharmaceutical composition comprising: (a) a binding protein of  claim 1 ,  3 ,  5 ,  10 ,  15 ,  21 ,  27 ,  36 ,  37 ,  38 , or  39 ; and (b) a pharmaceutically acceptable formulation agent. 
     
     
         51 . The pharmaceutical composition of  claim 50 , wherein the pharmaceutically acceptable formulation agent is a carrier, adjuvant, solubilizer, stabilizer, or anti-oxidant. 
     
     
         52 . A method for treating a metabolic disorder comprising administering to a human patient in need thereof the pharmaceutical composition of  claim 50 . 
     
     
         53 . The method of  claim 52 , wherein the metabolic disorder is one or more of: diabetes, obesity, dyslipidimia, hypertension, hepatosteaotosis, non-alcoholic steatohepatitis (NASH), acute pancreatitis, metabolic syndrome, cardiovascular disease, such as atherosclerosis, and aging. 
     
     
         54 . The method of  claim 53 , wherein the metabolic disorder is diabetes. 
     
     
         55 . The method of  claim 53 , wherein the metabolic disorder is obesity.

Cited by (0)

No later patents cite this yet.

References (0)

No backward citations on record.